The lion sprang ___ the cow:

This question assesses the correct usage of prepositions of movement. The verb "sprang" denotes a sudden, forceful leap.

• "upon" is the correct choice because it precisely conveys movement to a higher position on something, often implying impact or landing. It accurately describes the lion leaping onto the cow.
• "on" is a distractor; while sometimes interchangeable, "upon" often carries a slightly more formal or emphatic nuance for such dynamic actions.
• "over" is incorrect as it suggests movement above something, typically without making contact, or crossing from one side to another. The lion landed on the cow, not merely passed above it.
• "into" is incorrect as it signifies movement inside an enclosed space, which is not relevant to the context of a lion attacking a cow.